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The Fourth Doctor found that K9 had contracted laryngitis and had lost his voice as a result of this. K9 was left in the TARDIS while the Doctor and Romana II battled the Daleks and Davros. (TV: Destiny of the Daleks) During a later adventure, he remained patiently in the TARDIS, which would imply that his voice had not yet returned. (TV: City of Death) After his voice returned, it sounded quite different and remained so for a while (TV: The Creature from the Pit) before the familiar voice returned. (TV: The Leisure Hive)
K9 Mark 2 was the result of the activation of the Mark I's onboard regeneration unit. He had only very limited memories of his past as the Mark I. He had many adventures with Alistair Gryffen, Jorjie Turner, Darius Pike and his new master Starkey. (TV: Regeneration, The Bounty Hunter) Orthrus was an attempt by Drake and the Department to replicate K9, but it was of vastly inferior design. (TV: Jaws of Orthrus)
While caught up in a Time war between the past and the future on the planet Aoris, K9 was forced to spend a thousand years hidden on the planet, setting himself up as 'the great Machina' so that he could found a cult that would aid the Doctor and Romana in bringing the war to an end, hiding behind a fake computer with a version of the TARDIS until he could return to the Doctor and Romana once the war ended (AUDIO: The Paradox Planet, Legacy of Death).
Introduced into the plot in The Invasion of Time, K9 Mark II first actually appear on-screen in The Ribos Operation (1978). More mobile than his predecessor, Mark II exhibited the ability to sense and warn others of danger. He travels as a companion alongside the Doctor and Romana. Around the time of Romana's regeneration (the transition from actress Mary Tamm to Lalla Ward), the K9 character was explained as suffering from "laryngitis" to accommodate Leeson's departure from the series at the start of the 1979–80 season; for this time, he was portrayed by David Brierley, until Leeson's return for the 1980–81 season. When the Doctor and Romana travel to the parallel universe of E-Space, K9 is severely damaged, in Warriors' Gate (1981). The damage was such that K9 could only function in E-Space; when Romana decided to stay and forge her own path, the Doctor gave K9 to her. The character makes subsequent appearances in remakes of the unfinished serial Shada, alongside the Eighth Doctor (Paul McGann); the 2003 Shada audio play and webcast depict K9 Mark II as having returned to Gallifrey with Romana, now Lady President of the Time Lords, where the Doctor visits them. It is not clear if either of these audio appearances are considered canonical in relation to the continuity of the TV series.